| Latin and Greek DerivationsMany of these combining forms may be used as either  prefixes or suffixes. They are listed here, in alphabetical order, first those most commonly used as prefixes, then those commonly used as suffixes. Examples show current usage. 
          
            | Prefixes | Derived From | Meaning | Example |  
            | 
              a-, ab- | Latin | off, from, down, away | abduct, avert |  
            | 
              a-, an- | Greek    | not, without, less | abiotic, anaerobic |  
            | 
              actin- | Greek aktis | a ray, beam, spoke | actin, actinomycete |  
            | 
              ad- | Latin | to, attached to | adsorption |  
            | 
              aer- | Greek | air | aerobic |  
            | 
              amphi- | Greek | both, about, around | amphibian |  
            | 
              ana- | Latin | away, through, again | analysis |  
            | 
              andro- | Greek | man, male | androgens |  
            | 
              angio- | Greek | a vessel, closed    container | angiospermae |  
            | 
              anthropo- | Greek | referring to man | anthropology |  
            | 
              ant-, anti- | Greek | against, away, opposite | antibiosis |  
            | 
              ante- | Latin | before | anteroom |  
            | 
              ap-, aph-, apo- | Latin | from, off, separate | apogee |  
            | 
              aqua- | Latin | water | aquatic |  
            | 
              arche-, archeo- | Greek | ancient, primitive | archeology |  
            | 
              arthri-, arthro- | Greek arthron | joint, jointed | arthritis, arthrospore |  
            | 
              asco- | Greek askos | bag, sack, bladder | ascospore |  
            | 
              aureo- | Latin aureus | gold coloured | aureomycin |  
            | 
              auto- | Greek autos | self | autoimmune |    
          
            | Prefixes | Derived From | Meaning | Example |  
            | 
              bi- | Latin | two, twice, double | bipolar, binocular |  
            | 
              bio-, bios- | Greek | related to life | biology, biocidal |  
            | 
              blasto- | Greek blastos | an embryonic layer or    cell | blastomere |  
            | 
              brachy- | Greek | short | brachycephalic |  
            | 
              brad-, brady- | Greek | slow, slowness | bradycardia |  
            | 
              bry-, bryo- | Greek bryon | moss, mossy | bryophyte |  
            | 
              calic-, calix- | Latin | cup-like | calyx |  
            | 
              cani-, canis- | Latin | dog | canine |  
            | 
              cardia- | Greek kardia | heart | cardiac |  
            | 
              carn- | Latin carnis | flesh | carnivore |  
            | 
              carp- | Latin carpalis | wrist, bones | carpel |  
            | 
              cata- | Greek | decomposition,    degradation | catabolism |  
            | 
              cell- | Latin cella | small room | cellular |  
            | 
              cephal- | Latin | head | cephalic |  
            | 
              chloro- | Greek chloros | green, containing    chloride | chlorophyll |  
            | 
              chroma-, chromo- | Greek | coloured | chromosome |  
            | 
              chron-, chrono | Greek chronos | time | chronometer |  
            | 
              circum- | Latin | around, near, about | circumnavigate |  
            | 
              coel- | Greek koilos | hollow cavity, belly | coelom |  
            | 
              col-, com-, con- | Latin | with, together | combine, collide |  
            | 
              contra- | Latin | against | contradict |  
            | 
              crypto- | Greek kryptos | hidden | cryptogamic |  
            | 
              cyano- | Greek kyanos | dark blue, blue-green | cyanobacteria |  
            | 
              cyst- | Greek kystis | bladder | cystitis, cystidium |  
            | 
              cyt-, cyte-, cyto- | Greek kytos | cell, a hollow vessel | cytology |  
            | 
              de- | Latin | undoing, removal of,    from | dehydration |  
            | 
              den-, dent-  | Latin dens  | tooth  | dentition  |  
            | 
              dendro-  | Greek | tree | dendrochronology |  
            | 
              derm-, derma- | Greek | skin, hide | dermatitis |  
            | 
              deut-, deutero-  | Greek  | second, secondary  | deuterium  |  
            | 
              di-  | Greek  | double, twice, two  | disaccharide  |  
            | 
              dia-  | Greek  | through, across  | diameter  |  
            | 
              diplo-  | Greek  | twofold, double  | diploid  |  
            | 
              dis-  | Latin  | apart, away  | dissolve  |  
            | 
              dorm-  | Latin  | to sleep  | dormant, dormitory  |  
            | 
              drom-, drome-  | Greek  | a running, racing  | dromedary  |    
          
            | Prefixes | Derived From | Meaning | Example |  
            | 
              e-, ec-  | Latin  | out, out of  | efferent  |  
            | 
              eco-  | Greek oikos  | house, environment  | ecology  |  
            | 
              ecto-  | Greek ektos  | outside  | ectoderm  |  
            | 
              en-, endo-  | Greek endon  | within, internal  | endoskeleton  |  
            | 
              entero- | Greek enteron  | intestine | enterocolitis |  
            | 
              entomo- | Greek entoma  | insect | entomology |  
            | 
              eo-, eos-  | Greek | the dawn  | Eocene, Eohippus  |  
            | 
              epi-  | Greek | upon, above, top  | epidermis  |  
            | 
              erythro-  | Greek | red  | erythrocyte  |  
            | 
              eu-  | Greek | proper, true, good  | Eukaryotic, eufungi  |  
            | 
              ex-  | Latin | out, from  | excise  |  
            | 
              exo- | Greek | outer, external | exoskeleton |  
            | 
              extra- | Latin exter          | outside of, beyond | extracellular |  
            | 
              flagell- | Latin flagrum | whip, whip-like | flagellum |  
            | 
              fuc-, fuco- | Greek phyktos  | seaweed, algae, lichen  | fucoxanthin  |  
            | 
              gamo- | Greek gamos | sexual union | gamogenesis |  
            | 
              gastero-, gastro- | Greek gaster | stomach, belly | gastroenteritis |  
            | 
              geno- | Latin gene | origin, development | genotype |  
            | 
              ge-, geo- | Greek | earth | geology |  
            | 
              glu-, glyco- | Greek | sweet, sugar | glucose, glycogen |  
            | 
              gon-, goni-, gono- | Greek | reproductive, sexual | gonorrhea |  
            | 
              gymn-, gymno- | Greek gymnos | naked, bare | gymnosperm |  
            | 
              gyn-, gyne-, gyno- | Greek | woman, female | gynecology |  
            | 
              halo- | Greek hals | salt | halophile |  
            | 
              haplo- | Greek haploos | single | haploid |  
            | 
              heme-, hemo- | Greek haimo | blood | haematologist |  
            | 
              hemi- | Greek | half | hemisphere |  
            | 
              hepta- | Greek | seven | heptane |  
            | 
              herb- | Latin herba | pertaining to plants | herbicide |  
            | 
              hetero- | Greek | different, other,    unlike | heterozygous |  
            | 
              hex-, hexa- | Greek | six | hexagonal |  
            | 
              hipp-, hippo- | Greek hippos | horse | hippodrome |  
            | 
              histo- | Greek histos | tissue | histology |  
            | 
              holo- | Greek holos | whole, entire | holoblastic |  
            | 
              homeo, homo- | Greek | same, similar, like | homogeneous |  
            | 
              hyal-, hyalo- | Greek hyalos | glassy, transparent | hyaloid |  
            | 
              hydr-, hydro- | Greek | pertaining to water | hydrolysis |  
            | 
              hyper- | Greek | above, more, over | hyperactive |  
            | 
              hypo- | Greek | below, less, under | hypodermic |    
          
            | Prefixes | Derived From | Meaning | Example |  
            | 
              ichthy-, ichthyo- | Greek | referring to fish | ichthyology |  
            | 
              inter- | Latin | between | intercellular |  
            | 
              intra- | Latin | within, inside | intracellular |  
            | 
              intro- | Latin | inward, within | introvert |  
            | 
              iso- | Greek | equal, same | isotonic |  
            | 
              kine- | Greek | movement, moving | kinetics |  
            | 
              leuc-, leuk- | Greek | white | leucocyte |  
            | 
              lycan- | Greek lykos | wolf | lycanthropy |  
            | 
              macro- | Greek | large, big, long | macromolecule |  
            | 
              man-, manu- | Latin | hand | manual |  
            | 
              mastig- | Greek mastigos | whip | mastigophora |  
            | 
              meg-, mega- | Greek | great, large | megabyte |  
            | 
              melan-,melano- | Greek | black, dark | melanin |  
            | 
              mero- | Greek merus | part, piece | meroblast |  
            | 
              mes-, meso- | Greek mesos | middle, in between | mesoderm |  
            | 
              met-, meta- | Greek | later, following,    changed in position or form | metamorphosis |  
            | 
              micro- | Greek mikros | small | microbiology |  
            | 
              milli- | Latin | a thousandth part | millimetre |  
            | 
              mio- | Greek meion | less, smaller | Miocene |  
            | 
              mito-  | Greek mitos | thread | mitosis |  
            | 
              mon-, mono- | Greek | one, single | monocular |  
            | 
              morph- | Greek | shape, form | morphology |  
            | 
              mor-, mort- | Latin | die, death | mortality |  
            | 
              muc-, muco- | Latin | consisting of many    units | multicellular |  
            | 
              mus- | Latin | mouse, as one running | muscle |  
            | 
              myco-, mykos- | Greek | fungus, mushroom | mycology |  
            | 
              myo- | Greek mys | muscle | myoglobin |  
            | 
              myxo- | Greek | slime, mucus | myxomycetes |  
            | 
              nemato- | Greek | thread, threadlike | nematode |  
            | 
              ob- | Latin | against | obtuse |  
            | 
              octa- | Greek | eight | octopus |  
            | 
              olig-, oligo- | Greek | few, small, less | oligarchy |  
            | 
              omni- | Latin | all, everywhere | omnipotent |  
            | 
              oo- | Greek | pertaining to an egg | oocyte |  
            | 
              ophthalmo- | Greek | referring to the eye | ophthalmologist |  
            | 
              opisth-, opistho- | Greek | behind, backwards, back | Opisthobranchia |  
            | 
              orni-, ornitho- | Greek | bird | ornithology |  
            | 
              orth-, ortho- | Greek | straight | orthodontist |  
            | 
              osteo- | Greek | bone | osteocyte |  
            | 
              oto- | Greek | referring to the ear | otology |  
            | 
              ova-, ovi-, ovul- | Latin | egg | ovary, oviduct |    
          
            | Prefixes | Derived From | Meaning | Example |  
            | 
              paleo- | Greek | old, ancient | paleontology |  
            | 
              para- | Greek | beside, near, beyond | parasitism |  
            | 
              path-, patho- | Greek | disease, suffer | pathogenic |  
            | 
              ped-, pedi- | Latin | foot | pedicure |  
            | 
              penna-, pinna- | Latin | feather, feathery | pinnate |  
            | 
              pent-, penta- | Greek | five | pentagon |  
            | 
              per- | Latin | through | pervade, peruse |  
            | 
              peri- | Greek | around, surrounding | perimeter |  
            | 
              pher- | Greek | bearing, carrying,    support | pheromone |  
            | 
              phil- philo- | Greek | loving, attracted to | philanthropy |  
            | 
              phob- | Greek | fear, fearing | phobic |  
            | 
              photo- | Greek | pertaining to light | photosynthesis |  
            | 
              phyco- | Greek | seaweed, algae | phycology |  
            | 
              phylo- | Greek | tribe, race, related    group | phylogeny |  
            | 
              phyto- | Greek | pertaining to plants | phytohormone |  
            | 
              plasm-, plasma- | Greek | formative substance | plasmablasts |  
            | 
              plati-, platy- | Greek | flat | platypus |  
            | 
              pleio-, pleo- | Greek | more, many | pleiomorphic |  
            | 
              pod-, poda-, podi- | Greek | foot | podiatrist |  
            | 
              poly- | Greek | many | polyhedron |  
            | 
              post- | Latin | after | postnatal |  
            | 
              pre- | Latin | before | prenatal |  
            | 
              preter- | Latin | beyond | preterhuman |  
            | 
              prim- | Latin | first | primary |  
            | 
              pro- | Greek | before, on behalf of | proboscis |  
            | 
              pro- | Latin | forward | progressive |  
            | 
              proto- | Greek | first, primary | protozoa |  
            | 
              pseudo-  | Greek  | false  | pseudopod  |  
            | 
              psilo-  | Greek  | bare, mere  | psilopsida  |  
            | 
              pteri-, ptero- | Greek | fern, feather | pteridophyte |  
            | 
              quadr-, quadri-  | Latin  | four  | quadruped  |  
            | 
              radi-  | Latin  | ray, spoke of wheel  | radial  |  
            | 
              re-  | Latin  | back, again  | repeat  |  
            | 
              retro-  | Latin  | backward  | retroactive  |  
            | 
              rhiz-, rhizo-  | Greek  | pertaining to roots  | rhizoids  |  
            | 
              rhod-, rhodo-  | Greek  | a rose, red  | rhodopsin  |  
            | 
              rota-  | Latin  | wheel  | rotate  |    
          
            | Prefixes | Derived From | Meaning | Example |  
            | 
              sapr-, sapro-  | Greek  | rotten, putrid, dead  | saprobe  |  
            | 
              sarc-, sarco-  | Greek  | flesh, fleshy  | sarcoma  |  
            | 
              schiz-, schizo- | Greek | split, splitting | schizophrenia |  
            | 
              se- | Latin | apart | secede |  
            | 
              semi- | Latin | half | semicircle |  
            | 
              soma-, somato- | Greek | body | somatic |  
            | 
              sperma-, spermato- | Greek | seed | spermatozoa |  
            | 
              sporo- | Greek | spore | sporophyte |  
            | 
              staphylo- | Greek | bunch of grapes | staphylococcus |  
            | 
              stoma- | Greek | mouth | stomata |  
            | 
              strepto- | Greek | twisted, string of | streptococcus |  
            | 
              sub- | Latin | below, under, smaller | subapical |  
            | 
              supra-, super- | Latin | above, over | supernova |  
            | 
              sym-, syn- | Greek | together, with | synthesis |  
            | 
              taxi-, taxo- | Greek | to make order,    arrangement | taxonomy |  
            | 
              tel-, tele-, telo- | Greek | distant, end | telophase |  
            | 
              terra-, terre- | Latin | land, earth | terrestrial |  
            | 
              tetra- | Greek | four | tetrapod |  
            | 
              therm-, thermo- | Greek | heat | thermometer |  
            | 
              thigmo- | Greek | touch | thigmotropism |  
            | 
              trans- | Latin | across, through, over | transfer |  
            | 
              tri- | Latin | three | triangle |  
            | 
              tricho- | Greek | hair | trichogyne |  
            | 
              triplo- | Latin | triple | triploid |  
            | 
              troche-, trocho- | Greek | wheel, hoop | trochophore |  
            | 
              tropho- | Greek | nourishment | trophoblast |  
            | 
              ultra- | Latin | beyond, exceedingly | ultraconservative |  
            | 
              uni- | Latin | consisting of one | unicellular |  
            | 
              vice- | Latin | in place of | vice-president |  
            | 
              vid-, vis- | Latin | see | vision |  
            | 
              xen-, xeno- | Greek | dry, desert | xerophyte |  
            | 
              zoo- | Greek | animal, life | zoology |  
            | 
              zyg-, zygo- | Greek | to join together | zygote |    
          
            | Suffixes | Derived From | Meaning | Example |  
            | 
              -biosis | Greek | mode of living, way of    life | symbiosis |  
            | 
              -blast | Greek | formative, embryonic | mesoblast |  
            | 
              -chaeta-, -chete  | Greek  | a bristle  | Polychaeta  |  
            | 
              -chrome | Greek | colour | cytochrome |  
            | 
              -cidal, -cide | Latin | killer, a killing | fungicide |  
            | 
              -cocci, -coccus | Greek | round, seed, kernel | Streptococcus |  
            | 
              -cyst | Greek | pouch, sac | trichocyst |  
            | 
              -dactyl | Greek | finger | pentadactyl |  
            | 
              -derm, -dermis | Greek | skin, layer | epidermis |  
            | 
              -elle, -ule, -la, -le, -let, -ole | Latin  | small, diminutive    endings  | globule, piglet  |  
            | 
              -emia  | Greek  | blood disease  | anemia  |  
            | 
              -fer  | Latin  | bearer, producer, carry  | conifer, transfer  |  
            | 
              -gamous, -gamy  | Greek  | marriage, sexual fusion  | polygamy  |  
            | 
              -gen, -geny  | Greek  | origin, production  | progeny, hydrogen  |  
            | 
              -genesis  | Latin  | origin, development of  | embryogenesis  |  
            | 
              -gony  | Latin  | something produced  | cosmogeny  |  
            | 
              -graph  | Greek  | drawing, writing  | chromatograph  |  
            | 
              -hedral, -hedron | Greek | side | polyhedral |  
            | 
              -hydrate | Greek | compound of water with another    substance | carbohydrate |  
            | 
              -ism | Greek | act, practice or result    of | terrorism |  
            | 
              -ite | Latin | a division or part | somite |  
            | 
              -itis | Greek | inflammation or    infection | appendicitis |  
            | 
              -jugal, -jugate | Latin | to yoke, join together | conjugate |  
            | 
              -logy | Greek logos | science or study of | biology |  
            | 
              -lysis, -lytic | Greek | loosening, separation, splitting    into smaller units | photolysis |  
            | 
              -mer, -merous | Greek meros | a part, piece | polymer |  
            | 
              -meter | Greek metron | a measurement | diameter |  
            | 
              -morph | Greek | form | endomorph |  
            | 
              -mycin | Greek | derived from a fungus | aureomycin |  
            | 
              -nomy | Greek | systematized knowledge    of | astronomy |  
            | 
              -oma | Greek | tumorous | carcinoma |  
            | 
              -osis, -otic | Greek | abnormal condition,    disease | neurosis |  
            | 
              -phage | Greek | eater | bacteriophage |  
            | 
              -phase | Greek | a stage or condition | metaphase |  
            | 
              -phor, -phore | Greek | bearing, carrying, supporting | sporangiophore |  
            | 
              -phyll | Greek | leaf | chorophyll |  
            | 
              -phyta, -phyte | Greek | plant | epiphyte |  
            | 
              -plasm | Greek | formative substance | cytoplasm |  
            | 
              -plast | Greek | organized particle,    granule | choroplast |  
            | 
              -pod, -poda | Greek | foot | arthropod |  
            | 
              -some | Greek | body | chromosome |  
            | 
              -stasis | Greek | a stationary position | homeostasis |  
            | 
              -stat, -static | Greek | stationary, still | thermostat |  
            | 
              -stomy | Greek | opening into | colostomy |  
            | 
              -therm | Greek | heat | homoeotherm |  
            | 
              -thes, -thesis | Greek | arrangement, in order | hypothesis |  
            | 
              -tom, -tomy | Greek | dividing, surgery | lobotomy |  
            | 
              -trope, -tropic | Greek | turning | phototropic |  
            | 
              -vor, -vore | Latin vorare | feeding | carnivore |  
            | 
              -zoa, -zoan, -zoic | Greek | animal, life | protozoa |  The letters of the Greek alphabet are also used as  symbols by   mathematicians and scientists. These people pronounce the names of the letters   quite differently from Greek people, the 'scientific pronunciation' being very loosely based on the pronunciation of   classical Greek. The table below shows the full Greek alphabet in capital (upper case) and lower case characters. The scientific pronunciation of the letters is shown in the column headed Name in 
  English; the name of the letter and its pronunciation in modern Greek is indicated in the other two columns.  Note that the capital letters B,   H, P, X and Y look the same as letters in the Roman alphabet used in English, but in Modern Greek they are pronounced very differently. 
        
          
            | Capital | Lower Case
 | Name in English
 | Name as Pronounced
 in   Greek
 | Modern   Pronunciation |  
            | Α | α | alpha | alfa | a as in bath |  
            | Β | β | beta | veeta | v |  
            | Γ | γ | gamma | ghamma | Before A or O, this is a sound   that does not occur in English, like a soft g, the voiced version of the kh   below. Before E, I or Y it is pronounced y as in yet. |  
            | Δ | δ | delta | dhelta | dh represents the voiced th   sound in this and that |  
            | Ε | ε | epsilon | epsilon | e as in pet |  
            | Ζ | ζ | zeta | zeeta | z |  
            | Η | η | eta | eeta | ee as in feet |  
            | Θ | θ | theta | theeta | th unvoiced as in thin and   thanks |  
            | Ι | ι | iota | yotta | ee as in feet; like y in yes   when before vowel |  
            | Κ | κ | kappa | kappa | k |  
            | Λ | λ | lambda | lamdha | l |  
            | Μ | μ | mu | mee | m |  
            | Ν | ν | nu | nee | n |  
            | Ξ | ξ | xi | ksee | ks as x in fox, never as x in   xylophone |  
            | Ο | ο | omicron | omicron | o as in got |  
            | Π | π | pi | pee | p |  
            | Ρ | ρ | rho | ro | r trilled as in Spanish or   Italian |  
            | Σ | σ or ς | sigma | sighma | s unvoiced as in sauce, not   vase. The first lower-case form is for the start or the middle of words, the   second for the last letter of words. |  
            | Τ | τ | tau | taf | t |  
            | Υ | υ | upsilon | eepsilon | ee as in feet |  
            | Φ | φ | phi | fee | f |  
            | Χ | χ | chi | khee | kh represents ch sound in   Scottish word loch |  
            | Ψ | ψ | psi | psee | ps as in copse. The p is   pronounced even at the start of words. |  
            | Ω | ω | omega | omegha | o as in got |  
            |  |  |